Recovery and Management
of Peregrine Falcons in Virginia
Introduction
The
peregrine falcon (Falco
peregrinus) is a medium sized raptor with long pointed wings
and a long, narrow tail. The adult's plumage consists of a black
crown and nape, black "sideburns" and a spotted belly.
Peregrines are distributed essentially worldwide.
Three subspecies have been described in North America:
Peale's peregrine (F. p.
pealei) is the largest and darkest in color, and inhabits
the island chains of the Pacific Northwest; the Arctic peregrine
(F. p. tundrius) is a
small, pale, highly migratory form distributed in the Nearctic
region (most peregrines encountered on the eastern coast in the
fall and winter are of this subspecies); and the American
peregrine (F. p. anatum) is
intermediate in size, color, and migratory behavior.
Its range extends across the continent, reaching that of tundrius
in the north and extending south to north-central Mexico. The peregrines that originally inhabited western Virginia
were of an anatum subpopulation
called the Appalachian peregrine.
Those birds were larger and darker than others of the
subspecies, but that population became extinct before any
recovery efforts began.
Although
never occurring in large numbers, the American peregrine falcon
was once found throughout the Appalachian Mountains of the
eastern United States. Optimum
nesting habitat consisted of high, long cliffs, usually
overlooking water. Johannes
Gabler in 1983 reported the existence of 24 historical cliff
eyries in Virginia. Nesting
was also documented at 2 sites along the Virginia coast. At both
of these sites, falcons were observed using tree nests made by
other birds.
In 1942, the
known nesting population east of the Mississippi was estimated
at around 350 pairs. The
decline of the eastern peregrine resulted from human disturbance
and, in particular, the widespread use of organochlorine
pesticides such as DDT after WWII.
Peregrine falcons were extirpated from Virginia as a
nesting species by the mid-1960's.
In
response to range-wide declines and eventual extirpation in the
eastern United States, the American and Arctic peregrine falcon
subspecies were listed as endangered by the U.S. Fish and
Wildlife Service in 1970. The
Arctic peregrine recovered sufficiently to be delisted in
October, 1994. The
American subspecies, however, remained on the federal list until
August, 1999. Because
falcons have not recovered as a nesting population in their
historic mountainous range within Virginia, however, they remain
listed as a Threatened Species under Virginia's Endangered
Species Act and regulations enacted by the Department of Game
and Inland Fisheries. As
the Commonwealth's primary wildlife agency, the Department of
Game and Inland Fisheries has regulatory authority and the lead
role for recovery and management of peregrine falcons in
Virginia.
Initial
Recovery Efforts
In
1975, an Eastern Peregrine Falcon Recovery Team was appointed to
develop a federal Recovery Plan, subsequently completed in 1979.
The main objective of the plan was "to restore a new
self-sustaining population of peregrine falcons in the eastern
United States." Several
actions were described to accomplish this objective, including
protecting and managing essential nesting, wintering, and
migration habitat; eliminating the environmental pollutants that
originally caused most of the population declines; protecting
peregrines through law enforcement; implementing an education
program to build public support for and understanding of
peregrines; and releasing captive-produced falcons into the
wild. To provide a source of young birds for release, a captive
breeding facility was started in 1970 at the Laboratory of
Ornithology at Cornell University, and the first young falcons
were produced in 1973. The
breeding birds used were of a variety of subspecies from
different geographic areas including Spain, Scotland, Chile, the
Canadian tundra and taiga, the Alaska tundra and taiga, the
Aleutian Islands, the Queen Charlotte Islands, and California. Subsequent peregrine reintroduction in the eastern U.S. began
in 1974.
The
primary method used to reintroduce falcons to the wild is called
"hacking." This
process entails placing three to six birds, at approximately
28-30 days of age, into a protective box at the release site.
Food is provided daily in the form of domestic quail or
chicken. The hack
box is opened and the young falcons are released when they are
capable of flight (fledging), at approximately 40-45 days of
age. Food is
provided until the birds are able to successfully hunt.
The hope is that the young falcons will imprint on the
area and return as breeding adults in 2 to 3 years.
All released birds are individually marked with aluminum
U.S. Fish and Wildlife Service bands, and sometimes birds are
also marked with color-coded bands, or fitted with radio
transmitters.
Past
Recovery Efforts in Eastern VA
Efforts
to restore the peregrine in Virginia began in 1978, when 5 young
birds were released on Cobb Island in Northampton County.
Coastal release sites were initially chosen so that the
young birds could disperse in areas not typically inhabited by
great horned owls, known to be significant predators of young
falcons. These
coastal peregrines were to become part of a small, experimental
resident population. One
thought behind the coastal releases was that the young
peregrines would become self-sufficient in the vicinity of the
hack site and then wander extensively, returning as breeding
adults to natural cliff eyries of western Virginia.
Coastal
hacking continued until 1985.
In those 8 years a total of 115 birds were released from
8 sites, mostly on the Eastern Shore.
Some birds, however, were hacked at Back Bay National
Wildlife Refuge in Virginia Beach, and from the roof of a
nine-story building in Norfolk.
This urban site, first used in 1980, exhibited many of
the same qualities that made the marshes and islands ideal for
hacking. Those
qualities included relatively open terrain for hunting, abundant
prey items, minimal human disturbance, and protection from
predators. Over 85%
of the birds fledged from the coastal hack sites were reported
to have successfully dispersed after hacking.
Instead
of returning to the mountains to nest, however, the first
breeding pair of peregrines to become naturally established in
Virginia in more than 20 years took up residence in 1981 at the
hack tower on Wallops Island in Accomack County.
Similarly, the first successful nesting (3 fledglings)
occurred in 1982 at Assateague Island.
In subsequent years, pairs formed at all of the original
hack towers and other suitable sites, including an abandoned
shack and a retired ship in the James River reserve or
"ghost" fleet. In
the late 1980's, birds began appearing at some of the major
bridges on the coastal plain. The most productive bridge site has been the James River
Bridge in Newport News. Unfortunately,
birds hatched on bridges tend to have low survival rates, due to
the tenuous nest sites often chosen.
Cross-fostering, brood manipulation, and placement of
nest boxes are necessary to improve survival at these locations.
In
addition to young bridge birds being killed after falling from
the nest, there are other factors that increase chick mortality
and depress nest productivity at the coastal sites.
Despite early belief that the islands would be void of
great homed owls, predation and the displacement of adult
peregrines by great homed owls became a significant problem in
the mid-1990's. Placing
owl excluder bars on the nest box openings has reduced but not
eliminated this problem. Also,
human disturbance of some nests has occurred over the years.
Finally,
despite establishment of a nesting coastal population, some eggs
fail to hatch. Analysis
of some “addled” eggs has revealed high levels of DDE (a
byproduct of DDT) and other contaminants.
Although use of such pesticides as DDT has been banned in
the U.S., they are still widely used in other countries.
The highly migratory behavior of peregrines and many of
their prey species continues to expose peregrine falcons to
these substances.
Past
Recovery Efforts in Western VA
In
1983, Johannes Gabler surveyed 13 of the 24 known historical
peregrine nest sites in the mountains to locate any resident
peregrines, to determine the suitability of the sites for
releasing birds, and to describe the characteristics of the
historic eyries. Acknowledging
difficulty in verifying some of the historic sites, Gabler
nonetheless found no peregrines or any evidence of breeding.
He did, however, rate the suitability of each site he
visited for releasing captive produced falcons, and described
his reservations about using particular sites.
Subsequently,
the first mountain hack sites were established in 1985 on Mt.
Rogers and Clinch Mountain in southwest Virginia.
In 1988, two more sites were established in the George
Washington and Jefferson National Forests.
Falcons were hacked at sites in Shenandoah National Park
starting in 1989. The
only urban hack site in western Virginia was in downtown Roanoke
in 1992 and 1993. All
told, 131 peregrines were released from 9 western Virginia sites
from 1985 through 1993, with a 90% success rate.
In
1990, project personnel and volunteers, coordinated by the U.S.
Forest Service and National Park Service, began extensive cliff
monitoring. The
survey was initiated to search for peregrine activity and to
further evaluate cliffs as possible hack sites.
The surveys resulted in several records of single birds and
one pair of subadults being documented in Shenandoah County, and
also produced documentation of the first confirmed pairs in
Shenandoah Park. In
general, however, the surveys revealed few birds, and many
volunteers were uninspired to continue the effort after repeated
disappointment. Agency
biologists and cooperators continue to monitor some sites, but
there have been no widespread, formal surveys for peregrine
falcons in Virginia's mountains in recent years.
In
April of 1991, biologists from the Virginia Department of Game and
Inland Fisheries (DGIF), George Washington and Jefferson National
Forests (GWJNF) and the Center for Conservation Biology (CCB) of
The College of William & Mary participated in a helicopter
survey of historic and potential peregrine nest sites in
southwestern Virginia. The
two half-day flights covered approximately 26 locations. Sites were classified into four categories (A-D) with A being the most
promising sites, based on a variety of criteria including
vegetation around the cliff face, accessibility to humans and
mammalian predators, presence of sheltered holes and ledges, and
the presence of vultures, ravens, and/or owls.
No peregrines were seen during these surveys, although Dr.
Mitchell Byrd of the CCB had earlier observed a failed nesting
attempt at one of the sites examined.
Helicopter
surveys again were conducted in May of 1992 on the George
Washington National Forest and Shenandoah Park.
Approximately 100 cliffs were mapped as potentially good
nesting sites. Through
these surveys, photographs, and groundwork, Dr. Byrd produced a
list of sites to search in the future for breeding pairs of
peregrines.
The
first “new” pair of peregrine falcons in Virginia's
mountains was confirmed in 1992 at Little Stoney Man Mountain in
Shenandoah National Park; and in 1993, two pairs were seen on Old
Rag and Stoney Man mountains.
The first successful nesting was in 1994, when 2 chicks
fledged from the Stoney Man nest. The Stoney Man pair represents the only documented
successfully nesting peregrines in western Virginia since recovery
efforts began. They
raised young in 1994, 1996, and 1997, but spring storms caused egg
washouts in 1995 and 1998. The
pair failed again in 1999, and no nesting attempt was reported in
2000 or 2001.
2000
Efforts to Restore the Peregrine Falcon
In
the spring of 2000, neither member of the Stoney Man Mountain pair
of falcons returned to that site, and targeted surveys failed to
locate any peregrines in the portion of Shenandoah Park
encompassing Stoney Man, Hawksbill, and Old Rag mountains.
Thus, while the coastal population continued to be
productive, no peregrines pairs were known to exist in
Virginia's mountains. Considering
the normally poor survival of birds hatched on bridges, the
cooperating partners decided that some of those young could be
relocated and hacked into suitable mountain sites; a move intended
not only to reintroduce more birds to the mountains, but also to
enhance those birds' chances of survival.
Four young were removed from a nest on the James River
Bridge and moved to a hack box on Hawksbill Mountain.
They were released from the box on May 26; and that same
day, four more young from another bridge site were moved to a
temporary holding box at the same site.
Those young were soon moved to the hack box and released on
June 8. At that time,
all of the original four young had begun to disperse and hunt for
themselves, but they still were returning to the release site
occasionally to feed on quail being left by park employees for all
8 young birds. As of
June 16, all 8 had been observed flying, with some returning to
the release site carrying wild-captured prey.
2000
also was a hallmark year for introduction of peregrine falcons to
downtown Richmond, and marked the birth of this newest interagency
effort. In early May,
Dominion (formerly Virginia Power), proposed to hack falcons on
the roof of their 21-story corporate headquarters in downtown
Richmond. Funded by
Dominion, the initial effort was a cooperative venture among
Dominion, the Department of Game and Inland Fisheries, and The
Center for Conservation Biology at The College of William &
Mary. The first young
falcons released in Richmond were purchased from a captive breeder
in Idaho.
On
May 30, the 5 young falcons (4 males and 1 female) arrived at
Dulles Airport near Washington D.C.
They were received and transported by the U.S. Fish and
Wildlife Service to Richmond.
Local media covered placement of the birds into the hack
box and Dominion sponsored a contest to name them.
Once in the box, the young birds could be watched over the
Internet through a remote video camera.
When ready to fledge, the falcons were released and almost
immediately took to the air and disappeared from sight.
Because of their rather abrupt departure, biologists
initially were concerned for the young birds' welfare, but within
a few days all five had been observed near the release site.
FalconTrak
and Future Plans
Virginia's
coastal population of falcons seems to be relatively stable, if
small. One limiting
factor to population growth is the availability of nesting
structures. The
existing boxes and towers are repaired or replaced as resources
allow, and new sites are being created.
In 2000, boxes were placed on four power plant stacks in
eastern Virginia, in part because of successful nesting by
peregrines on such stacks in the Midwest.
Our
current objectives are to conduct rigorous surveys for nesting
pairs in Virginia's mountains, maintain and increase the coastal
population, focus on establishment/recovery of a mountain breeding
population, and answer basic questions regarding annual movements
of these highly migratory birds.
With over 100 birds released in Virginia's mountains
since 1985, it is likely that there are yet-undiscovered nesting
attempts occurring annually in Virginia's mountains.
Researchers who have been involved with the project for
decades tend to believe the birds are there; they just haven't
been found. Our
tripartite approach, namely (1) continuing to monitor and enhance
the survival of the coastal breeding population in Virginia; (2)
translocating and monitoring, through satellite telemetry, birds
produced on precarious coastal nests; and (3) initiating a
cooperative interagency/volunteer mountain nest survey protocol
offers the best opportunity in many years for recovery of
Virginia's most majestic falcon.
These efforts are intricately linked to our FalconTrak
partnership, now in our second year of operation.
For the latest
information, please see the
McGuireWoods
falcon site. |